By Dr Robert Arthur Lloyd

Journal articles

Martinac AD; Waters S; Lloyd RA; Bilston LE, 2026, 'Biomechanics of contagious yawning: Insights into cranio-cervical fluid dynamics and kinematic consistency', Respiratory Physiology and Neurobiology, 343, pp. 104575, http://dx.doi.org/10.1016/j.resp.2026.104575

Lloyd RA; Fletcher DF; Bilston LE, 2025, 'How volume changes in the epidural space drives respiratory cerebrospinal fluid flow', Journal of Biomechanics, 193, pp. 112983 - 112983, http://dx.doi.org/10.1016/j.jbiomech.2025.112983

Lloyd RA; Martinac AD; Liu Z; Berliner JA; Stoodley MA; Bilston LE, 2025, 'Chiari malformation does not obstruct cerebrospinal fluid flow during coughs', Computers in Biology and Medicine, 198, pp. 111208 - 111208, http://dx.doi.org/10.1016/j.compbiomed.2025.111208

Zhang Y; Lloyd R; Herbert R; Bilston L; Bolsterlee B, 2025, 'Prediction of Rotator Cuff Muscle Fibre Orientations Using a Population-Averaged Atlas Generated With Anatomical and Diffusion-Weighted Magnetic Resonance Images', NMR in Biomedicine, 38, pp. e70119, http://dx.doi.org/10.1002/nbm.70119

Lloyd RA; Dissanayake E; Jugé L; Bolsterlee B; Ball IK; Bilston LE, 2025, 'How mandibular and hyoid morphology alters tongue muscle architecture in healthy adults: An anatomical atlas and statistical shape model of the tongue', Computers in Biology and Medicine, 189, pp. 110006 - 110006, http://dx.doi.org/10.1016/j.compbiomed.2025.110006

Bolsterlee B; Lloyd R; Bilston LE; Herbert RD, 2025, 'A mechanically consistent muscle model shows that the maximum force-generating capacity of muscles is influenced by optimal fascicle length and muscle shape', Journal of Biomechanics, 182, pp. 112584, http://dx.doi.org/10.1016/j.jbiomech.2025.112584

Zhang Y; Lloyd R; Herbert RD; Bilston LE; Bolsterlee B, 2025, 'Prediction of Rotator Cuff Muscle Fibre Orientations Using a Population‐Averaged Atlas Generated With Anatomical and Diffusion‐Weighted Magnetic Resonance Images', NMR in biomedicine, 38

Hatt A; Lloyd R; Bolsterlee B; Bilston LE, 2023, 'Strain-dependent shear properties of human adipose tissue in vivo', Journal of the Mechanical Behavior of Biomedical Materials, 143, pp. 105924, http://dx.doi.org/10.1016/j.jmbbm.2023.105924

Liu S; Bilston LE; Flores Rodriguez N; Wright C; McMullan S; Lloyd R; Stoodley MA; Hemley SJ, 2022, 'Changes in intrathoracic pressure, not arterial pulsations, exert the greatest effect on tracer influx in the spinal cord', Fluids and Barriers of the CNS, 19, pp. 14, http://dx.doi.org/10.1186/s12987-022-00310-6

Lloyd RA; Stoodley MA; Bilston LE, 2022, 'Statistical shape models of the posterior cranial fossa and hindbrain volumes may provide a more robust clinical metric for Chiari malformation', Journal of Biomechanics, 137, pp. 111093, http://dx.doi.org/10.1016/j.jbiomech.2022.111093

Babaei B; Fovargue D; Lloyd RA; Miller R; Jugé L; Kaplan M; Sinkus R; Nordsletten DA; Bilston LE, 2021, 'Magnetic Resonance Elastography Reconstruction for Anisotropic Tissues', Medical Image Analysis, 74, pp. 1 - 15, http://dx.doi.org/10.1016/j.media.2021.102212

Lloyd RA; Butler JE; Gandevia SC; Ball IK; Toson B; Stoodley MA; Bilston LE, 2020, 'Respiratory cerebrospinal fluid flow is driven by the thoracic and lumbar spinal pressures', Journal of Physiology, 598, pp. 5789 - 5805, http://dx.doi.org/10.1113/JP279458

Dawes BH; Lloyd RA; Rogers JM; Magnussen JS; Bilston LE; Stoodley MA, 2019, 'Cerebellar Tissue Strain in Chiari Malformation with Headache', World Neurosurgery, 130, pp. e74 - e81, http://dx.doi.org/10.1016/j.wneu.2019.05.211

Lloyd RA; Stoodley MA; Fletcher DF; Bilston LE, 2019, 'The effects of variation in the arterial pulse waveform on perivascular flow', Journal of Biomechanics, 90, pp. 65 - 70, http://dx.doi.org/10.1016/j.jbiomech.2019.04.030

Cirovic S; Lloyd R; Jovanovik J; Volk HA; Rusbridge C, 2018, 'Computer simulation of syringomyelia in dogs', BMC Veterinary Research, 14, http://dx.doi.org/10.1186/s12917-018-1410-7

Lloyd RA; Fletcher DF; Clarke EC; Bilston LE, 2017, 'Chiari malformation may increase perivascular cerebrospinal fluid flow into the spinal cord: A subject-specific computational modelling study', Journal of Biomechanics, 65, pp. 185 - 193, http://dx.doi.org/10.1016/j.jbiomech.2017.10.007

Mowlavi S; Engmann J; Burbidge A; Lloyd R; Hayoun P; Le Reverend B; Ramaioli M, 2016, 'In vivo observations and in vitro experiments on the oral phase of swallowing of Newtonian and shear-thinning liquids', Journal of Biomechanics, 49, pp. 3788 - 3795, http://dx.doi.org/10.1016/j.jbiomech.2016.10.011

Conference Papers

Lloyd R; Berliner J; Martinac A; Stoodley M; Bilston L, 2025, 'Using real-time phase contrast MRI to understand coughing associated headaches in patients with Chiari malformation', in Proceedings of the International Society for Magnetic Resonance in Medicine ... Scientific Meeting and Exhibition., International Society for Magnetic Resonance in Medicine (ISMRM), presented at 2025 ISMRM & ISMRT Annual Meeting, http://dx.doi.org/10.58530/2025/4727

Martinac A; Lloyd R; Waters S; Bilston L, 2023, 'Effect of yawning on CSF and blood flow through the neck', in Proceedings of the International Society for Magnetic Resonance in Medicine ... Scientific Meeting and Exhibition., International Society for Magnetic Resonance in Medicine (ISMRM), presented at 2023 ISMRM & ISMRT Annual Meeting, http://dx.doi.org/10.58530/2023/3182

Hatt A; Lloyd R; Bolsterlee B; Bilston L, 2022, 'Non-rigid registration of DIXON images is an accurate method for calculating physiological strain in adipose tissue', in Proceedings of the International Society for Magnetic Resonance in Medicine ... Scientific Meeting and Exhibition., International Society for Magnetic Resonance in Medicine (ISMRM), presented at Joint Annual Meeting ISMRM-ESMRMB ISMRT 31st Annual Meeting, http://dx.doi.org/10.58530/2022/4239

Lloyd R; Ball I; Bilston L, 2022, 'Using machine learning to build a population model of tongue muscle architecture based on mDIXON and diffusion tensor imaging', in Proceedings of the International Society for Magnetic Resonance in Medicine ... Scientific Meeting and Exhibition., International Society for Magnetic Resonance in Medicine (ISMRM), presented at Joint Annual Meeting ISMRM-ESMRMB ISMRT 31st Annual Meeting, http://dx.doi.org/10.58530/2022/0534

Conference Abstracts

Lloyd R; Jugé L; Bilston L, 2023, 'O043 Investigating Upper-Airway Mechanics with Computational Tongue Models', in SLEEP Advances, Oxford University Press (OUP), Adelaide, SA Australia, Vol. 4, pp. a16 - a17, presented at Sleep DownUnder 2023, Adelaide, SA Australia, 08 November 2023 - 11 November 2023, http://dx.doi.org/10.1093/sleepadvances/zpad035.043

Preprints

Martinac A; Waters S; Lloyd R; Bilston L, 2025, Biomechanics of yawning: insights into cranio-cervical fluid dynamics and kinematic consistency, http://dx.doi.org/10.2139/ssrn.5954157

Lloyd R; Fletcher D; Bilston L, 2025, How Volume Changes in the Epidural Space Drives Respiratory Cerebrospinal Fluid Flow, http://dx.doi.org/10.2139/ssrn.5318520

Zhang Y; Lloyd R; Herbert RD; Bilston L; Bolsterlee B, 2024, Prediction of Rotator Cuff Muscle Fibre Orientations Using a Population-Averaged Atlas Generated with Anatomical and Diffusion-Weighted Magnetic Resonance Images, http://dx.doi.org/10.2139/ssrn.4984830

Martinac AD; Waters S; Lloyd RA; Bilston LE, Biomechanics of yawning: insights into cranio-cervical fluid dynamics and kinematic consistency, http://dx.doi.org/10.64898/2025.12.17.695005

Lloyd RA; Martinac AD; Liu Z; Berliner JA; Stoodley MA; Bilston LE, Chiari malformation does not obstruct cerebrospinal fluid flow during coughs, http://dx.doi.org/10.1101/2025.05.01.25326708

Zhang Y; Lloyd R; Herbert RD; Bilston LE; Bolsterlee B, Prediction of rotator cuff muscle fibre orientations using a population-averaged atlas generated with anatomical and diffusion-weighted magnetic resonance images, http://dx.doi.org/10.21203/rs.3.rs-4683327/v1

Other

Lloyd R, 2025, Human tongue muscle atlas, http://dx.doi.org/10.17605/OSF.IO/Z7U59

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