By Associate Professor Mark Holzer

Journal articles

Middleton JT; Holzer M; Closset I; Jones J; Brzezinski MA, 2026, 'High-Latitude Controls on the Silicon Isotope Composition of Silicic Acid in the South Pacific', Global Biogeochemical Cycles, 40, http://dx.doi.org/10.1029/2025GB009008

Brzezinski MA; Closset I; Holzer M; Jones JL, 2025, 'The Silicon Isotopic Composition of the North Pacific: Insights From US GEOTRACES Section GP15 and Modeling', Global Biogeochemical Cycles, 39, http://dx.doi.org/10.1029/2025GB008557

Millet B; de Lavergne C; Gray WR; Éthé C; Madec G; Holzer M; DeVries T; Gebbie G; Roche DM, 2025, 'Deep Ocean Ventilation: A Comparison Between a General Circulation Model and Data-Constrained Inverse Models', Journal of Advances in Modeling Earth Systems, 17, http://dx.doi.org/10.1029/2024MS004914

Pasquier B; Holzer M; Chamberlain MA, 2024, 'The biological and preformed carbon pumps in perpetually slower and warmer oceans', Biogeosciences, 21, pp. 3373 - 3400, http://dx.doi.org/10.5194/bg-21-3373-2024

Holzer M; DeVries T; Pasquier B, 2024, 'Atmospheric pCO2 Response to Stimulated Organic Carbon Export: Sensitivity Patterns and Timescales', Geophysical Research Letters, 51, http://dx.doi.org/10.1029/2024GL108462

John SG; Liang H; Pasquier B; Holzer M; Silva S, 2024, 'Biogeochemical Fluxes of Nickel in the Global Oceans Inferred From a Diagnostic Model', Global Biogeochemical Cycles, 38, http://dx.doi.org/10.1029/2023GB008018

Pasquier B; Holzer M; Chamberlain MA; Matear RJ; Bindoff NL, 2024, 'Deoxygenation and Its Drivers Analyzed in Steady State for Perpetually Slower and Warmer Oceans', Journal of Geophysical Research: Oceans, 129, http://dx.doi.org/10.1029/2024jc021043

Sulpis O; Trossman DS; Holzer M; Jeansson E; Lauvset SK; Middelburg JJ, 2023, 'Respiration Patterns in the Dark Ocean', Global Biogeochemical Cycles, 37, http://dx.doi.org/10.1029/2023GB007747

Pasquier B; Holzer M; Chamberlain MA; Matear RJ; Bindoff NL; Primeau FW, 2023, 'Optimal parameters for the ocean's nutrient, carbon, and oxygen cycles compensate for circulation biases but replumb the biological pump', Biogeosciences, 20, pp. 2985 - 3009, http://dx.doi.org/10.5194/bg-20-2985-2023

Gray WR; de Lavergne C; Jnglin Wills RC; Menviel L; Spence P; Holzer M; Kageyama M; Michel E, 2023, 'Poleward Shift in the Southern Hemisphere Westerly Winds Synchronous With the Deglacial Rise in CO2', Paleoceanography and Paleoclimatology, 38, http://dx.doi.org/10.1029/2023PA004666

John SG; Kelly RL; Bian X; Fu F; Smith MI; Lanning NT; Liang H; Pasquier B; Seelen EA; Holzer M; Wasylenki L; Conway TM; Fitzsimmons JN; Hutchins DA; Yang SC, 2022, 'The biogeochemical balance of oceanic nickel cycling', Nature Geoscience, 15, pp. 906 - 912, http://dx.doi.org/10.1038/s41561-022-01045-7

Holzer M; DeVries T, 2022, 'Source-Labeled Anthropogenic Carbon Reveals a Large Shift of Preindustrial Carbon From the Ocean to the Atmosphere', Global Biogeochemical Cycles, 36, http://dx.doi.org/10.1029/2022GB007405

Holzer M, 2022, 'The Fate of Oxygen in the Ocean and Its Sensitivity to Local Changes in Biological Production', Journal of Geophysical Research Oceans, 127, http://dx.doi.org/10.1029/2022JC018802

Kwon EY; Holzer M; Timmermann A; Primeau F, 2022, 'Estimating Three-Dimensional Carbon-To-Phosphorus Stoichiometry of Exported Marine Organic Matter', Global Biogeochemical Cycles, 36, http://dx.doi.org/10.1029/2021GB007154

Holzer M; DeVries T; de Lavergne C, 2021, 'Diffusion controls the ventilation of a Pacific Shadow Zone above abyssal overturning', Nature Communications, 12, pp. 4348, http://dx.doi.org/10.1038/s41467-021-24648-x

Holzer M; Kwon EY; Pasquier B, 2021, 'A New Metric of the Biological Carbon Pump: Number of Pump Passages and Its Control on Atmospheric pCO2', Global Biogeochemical Cycles, 35, http://dx.doi.org/10.1029/2020GB006863

Holzer M; Chamberlain MA; Matear RJ, 2020, 'Climate-Driven Changes in the Ocean's Ventilation Pathways and Time Scales Diagnosed From Transport Matrices', Journal of Geophysical Research Oceans, 125, http://dx.doi.org/10.1029/2020JC016414

John SG; Liang H; Weber T; DeVries T; Primeau F; Moore K; Holzer M; Mahowald N; Gardner W; Mishonov A; Richardson MJ; Faugere Y; Taburet G, 2020, 'AWESOME OCIM: A simple, flexible, and powerful tool for modeling elemental cycling in the oceans', Chemical Geology, 533, pp. 119403, http://dx.doi.org/10.1016/j.chemgeo.2019.119403

Holzer M; Pasquier B; DeVries T; Brzezinski M, 2019, 'Diatom Physiology Controls Silicic Acid Leakage in Response to Iron Fertilization', Global Biogeochemical Cycles, 33, pp. 1631 - 1653, http://dx.doi.org/10.1029/2019GB006460

Holzer M; DeVries T; Smethie WJ, 2019, 'The ocean's global 39Ar distribution estimated with an ocean circulation inverse model', Geophys. Res. Lett., 46, pp. 7491 - 7499, http://dx.doi.org/10.1029/2019GL082663

Chamberlain MA; Matear RJ; Holzer M; Bi D; Marsland SJ, 2019, 'Transport matrices from standard ocean-model output and quantifying circulation response to climate change', Ocean Modelling, 135, pp. 1 - 13, http://dx.doi.org/10.1016/j.ocemod.2019.01.005

DeVries T; Holzer M, 2019, 'Radiocarbon and Helium Isotope Constraints on Deep Ocean Ventilation and Mantle-3He Sources', Journal of Geophysical Research: Oceans, 124, pp. 3036 - 3057, http://dx.doi.org/10.1029/2018jc014716

Pasquier B; Holzer M, 2018, 'The number of past and future regenerations of iron in the ocean and its intrinsic fertilization efficiency', Biogeosciences, 15, pp. 7177 - 7203, http://dx.doi.org/10.5194/bg-15-7177-2018

Pasquier B; Holzer M, 2018, 'Iron fertilization efficiency and the number of past and future regenerations of iron in the ocean', , http://dx.doi.org/10.5194/bg-2018-379

Holzer M; Smethie WM; Ting YH, 2018, 'Ventilation of the Subtropical North Atlantic: Locations and Times of Last Ventilation Estimated Using Tracer Constraints From GEOTRACES Section GA03', Journal of Geophysical Research Oceans, 123, pp. 2332 - 2352, http://dx.doi.org/10.1002/2017JC013698

Yang H; Waugh DW; Holzer M, 2018, 'Decoupling the Effects of Transport and Chemical Loss on Tropospheric Composition: A Model Study of Path-Dependent Lifetimes', Journal of Geophysical Research Atmospheres, 123, pp. 2320 - 2335, http://dx.doi.org/10.1002/2017jd027871

Pasquier B; Holzer M, 2017, 'Inverse-model estimates of the ocean’s coupled phosphorus, silicon, and iron cycles', Biogeosciences, 14, pp. 4125 - 4159, http://dx.doi.org/10.5194/bg-14-4125-2017

Ting YH; Holzer M, 2017, 'Decadal changes in Southern Ocean ventilation inferred from deconvolutions of repeat hydrographies', Geophysical Research Letters, 44, pp. 5655 - 5664, http://dx.doi.org/10.1002/2017GL073788

Pasquier B; Holzer M, 2017, 'Inverse-model estimates of the ocean's coupled phosphorus, silicon, and iron cycles', , http://dx.doi.org/10.5194/bg-2017-122

DeVries T; Holzer M; Primeau F, 2017, 'Recent increase in oceanic carbon uptake driven by weaker upper-ocean overturning', Nature, 542, pp. 215 - 218, http://dx.doi.org/10.1038/nature21068

Holzer M; DeVries T; Bianchi D; Newton R; Schlosser P; Winckler G, 2017, 'Objective estimates of mantle 3He in the ocean and implications for constraining the deep ocean circulation', Earth and Planetary Science Letters, 458, pp. 305 - 314, http://dx.doi.org/10.1016/j.epsl.2016.10.054

Holzer M; Frants M; Pasquier B, 2016, 'The age of iron and iron source attribution in the ocean', Global Biogeochemical Cycles, 30, pp. 1454 - 1474, http://dx.doi.org/10.1002/2016GB005418

Pasquier B; Holzer M, 2016, 'The plumbing of the global biological pump: Efficiency control through leaks, pathways, and time scales', Journal of Geophysical Research Oceans, 121, pp. 6367 - 6388, http://dx.doi.org/10.1002/2016JC011821

Frants M; Holzer M; DeVries T; Matear R, 2016, 'Constraints on the global marine iron cycle from a simple inverse model', Journal of Geophysical Research Biogeosciences, 121, pp. 28 - 51, http://dx.doi.org/10.1002/2015JG003111

Holzer M; Waugh DW, 2015, 'Interhemispheric transit time distributions and path-dependent lifetimes constrained by measurements of SF6, CFCs, and CFC replacements', Geophysical Research Letters, 42, pp. 4581 - 4589, http://dx.doi.org/10.1002/2015GL064172

Tisdell CC; Holzer M, 2015, 'Analysis of the boundary value problem associated with the nonrelativistic Thomas-Fermi equation for heavy atoms in intense magnetic fields', Differential Equations and Applications, 7, pp. 27 - 41, http://dx.doi.org/10.7153/dea-07-03

Orbe C; Newman PA; Waugh DW; Holzer M; Oman LD; Li F; Polvani LM, 2015, 'Air-mass origin in the arctic. Part II: Response to increases in greenhouse gases', Journal of Climate, 28, pp. 9105 - 9120, http://dx.doi.org/10.1175/JCLI-D-15-0296.1

Orbe C; Newman PA; Waugh DW; Holzer M; Oman LD; Li F; Polvani LM, 2015, 'Airmass origin in the Arctic. Part I: Seasonality', Journal of Climate, 28, pp. 4997 - 5014, http://dx.doi.org/10.1175/JCLI-D-14-00720.1

Holzer M; Brzezinski MA, 2015, 'Controls on the silicon isotope distribution in the ocean: New diagnostics from a data-constrained model', Global Biogeochemical Cycles, 29, pp. 267 - 287, http://dx.doi.org/10.1002/2014GB004967

Orbe C; Holzer M; Polvani LM; Waugh DW; Li F; Oman LD; Newman PA, 2014, 'Seasonal ventilation of the stratosphere: Robust diagnostics from one-way flux distributions', Journal of Geophysical Research, 119, pp. 293 - 306, http://dx.doi.org/10.1002/2013JD020213

Holzer M; Primeau FW; DeVries T; Matear R, 2014, 'The Southern Ocean silicon trap: Data-constrained estimates of regenerated silicic acid, trapping efficiencies, and global transport paths', Journal of Geophysical Research Oceans, 119, pp. 313 - 331, http://dx.doi.org/10.1002/2013JC009356

Holzer M; Polvani LM, 2013, 'Lifetime dependent flux into the lowermost stratosphere for idealized trace gases of surface origin', Journal of Geophysical Research, 118, pp. 9367 - 9375, http://dx.doi.org/10.1002/jgrd.50657

Orbe C; Holzer M; Polvani LM; Waugh DW, 2013, 'Air-mass origin as a diagnostic of tropospheric transport', Journal of Geophysical Research-Atmospheres, 118, pp. 1459 - 1470, http://dx.doi.org/10.1002/jgrd.50133

Holzer M; Primeau FW, 2013, 'Global teleconnections in the oceanic phosphorus cycle: Patterns, paths, and timescales', Journal of Geophysical Research, 118, pp. 1775 - 1796, http://dx.doi.org/10.1002/jgrc.20072

Waugh DW; Primeau FW; DeVries T; Holzer M, 2013, 'Recent changes in the ventilation of the southern oceans', Science, 339, pp. 568 - 570, http://dx.doi.org/10.1126/science.1225411

Primeau FW; Holzer M; DeVries T, 2013, 'Southern Ocean nutrient trapping and the efficiency of the biological pump', Journal of Geophysical Research, 118, pp. 2547 - 2564, http://dx.doi.org/10.1002/jgrc.20181

Orbe C; Holzer M; Polvani LM, 2012, 'Flux distributions as robust diagnostics of stratosphere-toposphere exchange', Journal of Geophysical Research, 117, pp. Article number: D01302, http://dx.doi.org/10.1029/2011JD016455

Holzer M; Orbe C; Primeau FW, 2012, 'Stratospheric mean residence time and mean age on the tropopause: Connections and implications for observational constraints', Journal of Geophysical Research-Atmospheres, 117, pp. D12314 (8pp), http://dx.doi.org/10.1029/2012JD017547

Khatiwala S; Primeau FW; Holzer M, 2012, 'Ventilation of the deep ocean constrained with tracer observations and implications for radiocarbon estimates of ideal mean age', Earth and Planetary Science Letters, 325-326, pp. 116 - 125, http://dx.doi.org/10.1016/j.epsl.2012.01.038

Holzer M; Primeau FW, 2010, 'Improved constraints on transit time distribtions from argon 39: A maximum entropy approach', Journal of Geophysical Research, 115, pp. c12021(19pp), http://dx.doi.org/10.1029/2010JC006410

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